A 58-year-old man from Mumbai presents with a 2-year history of progressive tremor at rest in his right hand, which worsens when walking. On examination, he has a characteristic 'pill-rolling' tremor, rigidity on passive movement of the wrist, and bradykinesia. His gait is shuffling with reduced arm swing. MRI brain is normal. Which of the following best explains the neurophysiological basis of his tremor?

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