A 67-year-old man undergoes robot-assisted radical prostatectomy for Gleason 4+3=7 (Grade Group 3) prostatic adenocarcinoma. The gross specimen is bivalved along the coronal plane. The structure marked A in the diagram shows a firm, gritty, yellow-tan tumour in the posterior peripheral zone. Which of the following pathological features BEST explains the firm and gritty texture of this lesion?

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