A 52-year-old woman presents with pulsatile tinnitus synchronous with her heartbeat in the right ear, progressive conductive hearing loss, and aural fullness over 8 months. Otoscopy reveals a reddish-blue pulsating retrotympanic mass in the inferior mesotympanum with positive Brown sign. High-resolution CT shows a soft tissue mass confined to the cochlear promontory without invasion of the jugular foramen. The structure marked **A** in the diagram represents this clinical presentation of glomus tympanicum with conductive loss amenable to surgical resection. Which of the following is the embryological origin of the paraganglioma cells that comprise this lesion?
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