A 28-year-old man presents with progressive facial dysmorphism, multiple lentigines on the lips and eyelids, and recent-onset headaches with visual field defects. Imaging reveals a growth hormone-secreting pituitary adenoma and an incidental cardiac myxoma in the right ventricle. Genetic testing identifies an inactivating mutation in the gene marked **A** on the diagram. Which of the following best explains the multi-organ tumorigenesis in this patient?
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