A 42-year-old Indian man presents with episodic severe headaches, profuse sweating, and palpitations for 3 months. Blood pressure is 165/105 mmHg during an episode and 145/90 mmHg at baseline. Imaging reveals a well-circumscribed, encapsulated adrenal medullary tumor (marked **A** in the diagram). Biochemical testing confirms elevated plasma-free metanephrines. Which of the following best describes the cellular origin and functional consequence of the structure marked **A**?
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